jueves, 14 de junio de 2007

¿¿¿WhAt's iS A DiNoSaUr????

WhAt'S iS a DiNoSaUr?
Definition


Triceratops skeleton at the American Museum of Natural History in New York City.
The taxon Dinosauria was formally named in 1840 by English palaeontologist Richard Owen, who used it to refer to the "distinct tribe or sub-order of Saurian Reptiles" that were then being recognized in England and around the world.[1] The term is derived from the Greek words δεινός (deinos meaning "terrible", "fearsome", or "formidable") and σαύρα (saura meaning "lizard" or "reptile"). Though the taxonomic name has often been interpreted as a reference to dinosaurs' teeth, claws, and other fearsome characteristics, Owen intended it merely to evoke their size and majesty.[2]
Dinosaurs were an extremely varied group of animals; according to a 2006 study, over 500 dinosaur genera have been identified with certainty so far, and the total number of genera preserved in the fossil record has been estimated at around 1,850, nearly 75% of which remain to be discovered.[3][4] An earlier study predicted that about 3,400 dinosaur genera existed, including many which would not have been preserved in the fossil record.[5] Some were herbivorous, others carnivorous. Some dinosaurs were bipeds, some were quadrupeds, and others, such as Ammosaurus and Iguanodon, could walk just as easily on two or four legs. Regardless of body type, nearly all known dinosaurs were well-adapted for a predominantly terrestrial, rather than aquatic or aerial, habitat.








Distinguishing features of dinosaurs
While recent discoveries have made it more difficult to present a universally agreed-upon list of dinosaurs' distinguishing features, nearly all dinosaurs discovered so far share certain modifications to the ancestral archosaurian skeleton. Although some later groups of dinosaurs featured further modified versions of these traits, they are considered typical across Dinosauria; the earliest dinosaurs had them and passed them on to all their descendants. Such common structures across a taxonomic group are called synapomorphies.
Dinosaur synapomorphies include an elongated crest on the humerus, or upper arm bone, to accommodate the attachment of deltopectoral muscles; a shelf at the rear of the ilium, or main hip bone; a tibia, or shin bone, featuring a broad lower edge and a flange pointing out and to the rear; and an ascending projection on the astragalus, one of the ankle bones, which secures it to the tibia.[6]


Edmontonia was an "armored dinosaur" of the group Ankylosauria.
A variety of other skeletal features were shared by many dinosaurs. However, because they were either common to other groups of archosaurs or were not present in all early dinosaurs, these features are not considered to be synapomorphies. Such shared features include a diapsid skull bearing two pairs of holes in the temporal region; holes in the snout and lower jaw (two characteristics shared by other archosaurs); loss of the skull's postfrontal bone; a long neck incorporating an S-shaped curve;[7] an elongated scapula, or shoulder blade; forelimbs shorter and lighter than hind limbs, coupled to asymmetrical hands; a sacrum composed of three or more fused vertebrae; and an acetabulum, or hip socket, with a hole at the center of its inside surface.[8]
The open, or "perforate", hip joint described above had significant implications for dinosaur movement and behavior. Most notably, it allowed dinosaur hind limbs to be "underslung", or situated directly beneath the animals' bodies; this, in turn, allowed dinosaurs to stand erect in a manner similar to modern mammals, but distinct from most other reptiles, whose limbs sprawl out to either side.[9] Vertical limb configuration also enabled dinosaurs to breathe easily while moving, which likely permitted stamina and activity levels that surpassed those of "sprawling" reptiles.















































Phylogenetic definition
Under phylogenetic taxonomy, dinosaurs are usually defined as all descendants of the most recent common ancestor of Triceratops and modern birds.[10] It has also been suggested that Dinosauria be defined as all the descendants of the most recent common ancestor of Megalosaurus and Iguanodon, because these were two of the three genera cited by Richard Owen when he recognized the Dinosauria.[11] They are divided into Ornithischia (bird-hipped) and Saurischia (lizard-hipped), depending upon pelvic structure. Ornithischian dinosaurs had a four-pronged pelvic configuration, incorporating a caudally-directed (rear-pointing) pubis bone with (most commonly) a forward-pointing process. By contrast, the pelvic structure of saurischian dinosaurs was three-pronged, and featured a pubis bone directed cranially, or forwards, only.[9] Ornithischia includes all taxa sharing a more recent common ancestor with Triceratops than with Saurischia, while Saurischia includes those taxa sharing a more recent common ancestor with birds than with Ornithischia.


Stegosaurus skeleton, Field Museum, Chicago.
There is an almost universal consensus among paleontologists that birds are the descendants of theropod dinosaurs. Using the strict cladistical definition that all descendants of a single common ancestor are related, modern birds are dinosaurs and dinosaurs are, therefore, not extinct. Modern birds are classified by most paleontologists as belonging to the subgroup Maniraptora, which are coelurosaurs, which are theropods, which are saurischians, which are dinosaurs.[12]
However, referring to birds as 'avian dinosaurs' and to all other dinosaurs as 'non-avian dinosaurs' is cumbersome. Birds are still referred to as birds, at least in popular usage and among ornithologists. It is also technically correct to refer to birds as a distinct group under the older Linnaean classification system, which accepts paraphyletic taxa that exclude some descendants of a single common ancestor. Paleontologists mostly use cladistics, which classifies birds as dinosaurs, but some biologists of the older generation do not.
For clarity, this article will use 'dinosaur' as a synonym for 'non-avian dinosaur', and 'bird' as a synonym for 'avian dinosaur' (meaning any animal that evolved from the common ancestor of Archaeopteryx and modern birds). The term 'non-avian dinosaur' will be used for emphasis as needed.
Size


Comparative size of Diplodocus; human figures provide scale.
While the evidence is incomplete, it is clear that, as a group, dinosaurs were large. Even by dinosaur standards, the sauropods were gigantic. For much of the dinosaur era, the smallest sauropods were larger than anything else in their habitat, and the largest were an order of magnitude more massive than anything else that has since walked the Earth. Giant prehistoric mammals such as the Indricotherium and the Columbian mammoth were dwarfed by the giant sauropods, and only a handful of modern aquatic animals approach or surpass them in size — most notably the blue whale, which reaches up to 190,000 kg (209 tons) and 33.5 m (110 ft) in length.
Most dinosaurs, however, were much smaller than the giant sauropods. Current evidence suggests that dinosaur average size varied through the Triassic, early Jurassic, late Jurassic and Cretaceous periods.[13] According to paleontologist Bill Erickson, estimates of median dinosaur weight range from 500 kg to 5 tonnes; a recent study of 63 dinosaur genera yielded an average weight greater than 850 kg — comparable to the weight of a grizzly bear — and a median weight of nearly 2 tons, or about as much as a giraffe. This contrasts sharply with the size of modern mammals; on average, mammals weigh only 863 grams, or about as much as a large rodent. The smallest dinosaur was bigger than two-thirds of all current mammals; the majority of dinosaurs were bigger than all but 2% of living mammals.[14]
Largest and smallest dinosaurs
Only a tiny percentage of animals ever fossilize, and most of these remain buried in the earth. Few of the specimens that are recovered are complete skeletons, and impressions of skin and other soft tissues are rare. Rebuilding a complete skeleton by comparing the size and morphology of bones to those of similar, better-known species is an inexact art, and reconstructing the muscles and other organs of the living animal is, at best, a process of educated guesswork. As a result, scientists will probably never be certain of the largest and smallest dinosaurs.


Comparative size of Sauroposeidon.


Comparative size of Eoraptor.
The tallest and heaviest dinosaur known from good skeletons is Brachiosaurus brancai (also known as Giraffatitan). Its remains were discovered in Tanzania between 1907–12. Bones from multiple similarly-sized individuals were incorporated into the skeleton now mounted and on display at the Humboldt Museum of Berlin;[15] this mount is 12 m (38 ft) tall, 22.5 m (74 ft) long, and would have belonged to an animal that weighed between 30,000–60,000 kg (33–66 short tons). The longest complete dinosaur is the 27 m (89 ft) long Diplodocus, which was discovered in Wyoming in the United States and displayed in Pittsburgh's Carnegie Natural History Museum in 1907.
There were larger dinosaurs, but knowledge of them is based entirely on a small number of fragmentary fossils. Most of the largest herbivorous specimens on record were all discovered in the 1970s or later, and include the massive Argentinosaurus, which may have weighed 80,000–100,000 kg (88–121 tons); the longest, the 40 m (130 ft) long Supersaurus; and the tallest, the 18 m (60 ft) Sauroposeidon, which could have reached a sixth-floor window. The longest of them all may have been Amphicoelias fragillimus, known only from a now lost partial vertebral neural arch described in 1878. Extrapolating from the illustration of this bone, the animal may have been 58 m (190 ft) long and weighed over 120,000 kg (132 tons),[16] heavier than all known dinosaurs except possibly the poorly known Bruhathkayosaurus, which could have weighed 175,000–220,000 kg (193–243 tons). The largest known carnivorous dinosaur was Spinosaurus, reaching a length of 16–18 meters (53–60 ft), and weighing in at 9 tons.[17] Other large meat-eaters included Giganotosaurus, Mapusaurus, Tyrannosaurus rex and Carcharodontosaurus.
Not including modern birds like the bee hummingbird, the smallest dinosaurs known were about the size of a crow or a chicken. The theropods Microraptor and Parvicursor were both under 60 cm (2 ft) in length.

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